Faculty

Caio Brunharo

Associate Professor of Plant Sciences

118 ASI
University Park, PA

Email brunharo@psu.edu
Phone 814-865-9552
Pronouns He/Him

Overview

Weed adaptation to agricultural practices and climate change; Weed biology, ecology, and evolution; Abiotic stress; Herbicide resistance.

Areas of Expertise

Herbicide Agricultural and Biological Sciences
Glyphosate Agricultural and Biological Sciences
Glyphosate Keyphrases
Herbicide Resistance Keyphrases
Herbicide Resistance Agricultural and Biological Sciences
Herbicide Keyphrases
Glyphosate Resistance Keyphrases
Genetics Agricultural and Biological Sciences

Sensitivity and resistance survey of common lambsquarters (Chenopodium album) populations to bentazon across the United States
Ivana Santos Moisinho, Ed Peachey, Martin M. Williams, Mark Vangessel, Barbara Scott, Lynn M. Sosnoskie, Pavle Pavlovic, Caio Brunharo, Jan 9 2026, In: Weed Science
Date: Jan 9 2026

Wash-off potential from living and aged cover crop residues differs among soil-residual herbicide sorption properties
Cody Smith, Caio Brunharo, Kyle Elkin, Michael Flessner, Mark Vangessel, John M. Wallace, 2026, In: Weed Science
Date: 2026

Detection and confirmation of atrazine-resistant Palmer amaranth (Amaranthus palmeri) in the northeastern United States
Thierry E. Besançon, Caio A. Brunharo, James J. Polashock, Lynn M. Sosnoskie, 2026, In: Weed Science
Date: 2026

YOLOv7-CBAM and DeepSORT with pixel grid analysis for Real-Time weed localization and Intra-Row density estimation in apple orchards
Lawrence Arthur, Sadjad Mahnan, Long He, Magni Hussain, Paul Heinemann, Caio Brunharo, Dec 2025, In: Computers and Electronics in Agriculture
Date: Dec 2025

Multiple Origins or Widespread Gene Flow in Agricultural Fields? Regional Population Genomics of Herbicide Resistance in Bromus tectorum
Victor H.V. Ribeiro, Joseph Gallagher, Carol Mallory-Smith, Judit Barroso, Caio A.C.G. Brunharo, Jun 2025, In: Molecular ecology
Date: Jun 2025

The Genome of Lolium multiflorum Reveals the Genetic Architecture of Paraquat Resistance
Caio A. Brunharo, Aidan W. Short, Lucas K. Bobadilla, Matthew A. Streisfeld, May 2025, In: Molecular ecology
Date: May 2025

Response of Horseweed (Erigeron canadensis) from New York Vineyards and Orchards to Paraquat and Diquat
Bethany M. Schulteis, Ivana Santos Moisinho, Aleah Butler-Jones, Thierry E. Besançon, Caio Brunharo, Lynn M. Sosnoskie, Apr 2025, In: HortScience
Date: Apr 2025

Real-time Weed Species Localization and Tracking for Intra-Row Weed Density Estimation in Apple Orchards
Lawrence Arthur, Sadjad Mahnan, Long He, Magni Hussain, Paul Heinemann, Caio Brunharo, 2025, In:
Date: 2025

Herbicide deposition and wash-off potential is affected by cover crop management tactics used in planting green systems
Cody Smith, Caio Brunharo, Kyle Elkin, Michael Flessner, Mark VanGessel, John M. Wallace, 2025, In: Weed Science
Date: 2025

Genetic diversity and population structure of ALS-resistant Amaranthus hybridus across Brazil's primary soybean-growing regions
Acácio Gonçalves Netto, Victor Hugo Vidal Ribeiro, Marcelo Nicolai, Ramiro Fernando Lopez Ovejero, Vanessa Francieli Vital Silva, Gilmar José Picoli Junior, Caio Brunharo, 2025, In: Pest Management Science
Date: 2025

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Review
Maor Matzrafi, Christopher Preston, Caio Augusto Brunharo, May 2021, In: Pest Management Science
Citations: 54

Role of Glutamine Synthetase Isogenes and Herbicide Metabolism in the Mechanism of Resistance to Glufosinate in Lolium perenne L. spp. multiflorum Biotypes from Oregon
Caio A.C.G. Brunharo, Hudson K. Takano, Carol A. Mallory-Smith, Franck E. Dayan, Bradley D. Hanson, Aug 7 2019, In: Journal of agricultural and food chemistry
Citations: 54

Non-target-Site Resistance in Lolium spp. Globally
Andréia K. Suzukawa, Lucas K. Bobadilla, Carol Mallory-Smith, Caio A.C.G. Brunharo, Jan 22 2021, In: Frontiers in Plant Science
Citations: 51

Confirmation and mechanism of glyphosate resistance in tall windmill grass (Chloris elata) from Brazil
Caio Acg Brunharo, Eric L. Patterson, Daniela R. Carrijo, Marcel Sc de Melo, Marcelo Nicolai, Todd A. Gaines, Scott J. Nissen, Pedro J. Christoffoleti, Sep 1 2016, In: Pest Management Science
Citations: 45

Increased temperatures and elevated CO <sub>2</sub> levels reduce the sensitivity of Conyza canadensis and Chenopodium album to glyphosate
Maor Matzrafi, Caio Brunharo, Parsa Tehranchian, Bradley D. Hanson, Marie Jasieniuk, Dec 1 2019, In: Scientific reports
Citations: 42

Vacuolar sequestration of paraquat is involved in the resistance mechanism in lolium perenne L. Spp. multiflorum
Caio A.C.G. Brunharo, Bradley D. Hanson, Aug 25 2017, In: Frontiers in Plant Science
Citations: 40

EPSPS duplication and mutation involved in glyphosate resistance in the allotetraploid weed species Poa annua L.
Caio Augusto De Castro Grossi Brunharo, Sarah Morran, Katie Martin, Marcelo L. Moretti, Bradley D. Hanson, Jun 2019, In: Pest Management Science
Citations: 29

Multiple Herbicide-Resistant Italian Ryegrass [Lolium perenne L. spp. multiflorum (Lam.) Husnot] in California Perennial Crops
Caio A.C.G. Brunharo, Bradley D. Hanson, Nov 1 2018, In: Weed Science
Citations: 27

Western United States and Canada perspective
Caio A.C.G. Brunharo, Roger Gast, Vipan Kumar, Carol A. Mallory-Smith, Breanne D. Tidemann, Hugh J. Beckie, 2022, In: Weed Science
Citations: 25

Population genomics of Digitaria insularis from soybean areas in Brazil
Acácio Gonçalves Netto, Erick M.G. Cordeiro, Marcelo Nicolai, Saul J.P. de Carvalho, Ramiro Fernando Lopez Ovejero, Caio A.C.G. Brunharo, Maria I. Zucchi, Pedro J. Christoffoleti, Dec 2021, In: Pest Management Science
Citations: 21

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