My research focuses on understand disease dynamics on a complex landscape. I study gray wolves in Yellowstone National Park, where multiple prey species, predators species, and pathogens interact. Some of my research questions include: What are the mechanisms underlying patterns of disease outbreak? How important is the wolf social structure in disease "trapping"? How does wolf disease impact trophic interactions? What are the drivers of the fission-fusion dynamics we observe in and between wolf packs?